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AUGUSTA, Ga. (AP) Animal welfare officials are accusing Georgia Regents University of performing unnecessary dental implant procedures on dogs, then euthanizing them.

Humane Society of the United States spokeswoman Stephanie Twining said Wednesday that a three-month undercover investigation revealed that dogs being used to test dental implants have had teeth pulled, replaced with implants, then are euthanized.

Twining says jaw bone samples were taken from dead dogs.

University Senior Vice President for Research Mark Hamrick said in a statement that researchers at the Augusta school use protocols are regularly reviewed by the United States Department of Agriculture, and that the university adheres to local, state and federal guidelines.

Hamrick says the school doesn't frequently use dogs for research and that all research projects involving animals are reviewed by the Institutional Animal Care & Use Committee.